An archesporial cell is a diploid sporophytic cell differentiated from the nucellar cells of the ovule of a flowering plant. This cell has the ability to form the female spore in later stages of the life cycle.

In angiosperms, the archesporial cell is differentiated from a sub-epidermal cell towards the micropylar end and undergoes a periclinal division to form a large Megaspore mother cell (MMC) and a primary sporogenous cell. The MMC further undergoes meiotic divisions to form the megaspores.